Denomination: Church of England
Division 1971); children– Lochlan Houghton, Gregg Bemis, Garth Anders. Bachelor of Arts, Macalester College, St. Paul, 1950. Doctor of Medicine Yale University, 1954.
Master of Arts, Cambridge University, 1964. Intern Johns Hopkins Hospital, Baltimore, 1954-1955, resident in pathology, 1955-1957. Visiting investigator Rockefeller Institute Medical Research, New York City, 1957-1960.
Postdoctoral fellow Karolinska Instituet, Stockholm, 1960-1961. Postdoctoral fellow Cambridge University, England, 1961-1964, lecturer in pathology, since 1964. Assistant editor Immunological Communications, since 1980.
Contributor articles to professional journals. National Institutes of Health fellow, 1959-1964.