Assistant professor biochemistry University Padova, University Pavia, Italy, 1953-1956. Postdoctoral fellow National Research Council, Ottawa, Canada, 1957-1959. Member staff Centre National de la Recherche Scientifique as chargé, 1959-1963, maître, 1963-1968, director research, since 1968.
Head laboratory molecular genetics Institute Jacques Monod, Paris, since 1969, director Institute, 1978-1981. Visiting science biophysics department Johns Hopkins University, Baltimore, 1962. Scholar Fogarty International Center, Bethesda, Maryland, 1981-1984.
Served with Italian Air Force, 1952-1953. Member Federation European Bio-chemical Societies (chairman advanced course committee 1978-1986, Diplome Honneur 1986), European Molecular Biology Organization, International Council of Scientific Unions (chairman committee on genetic experimentation since 1982).