Bachelor of Mathematics, University Montpellier, 1942. Doctor of Medicine, University Lyons, 1949. Dsc, Sorbonne, 1959. Médecin auxiliaire Medical Corps, French Army, 1943, advanced through grades to Commander.
Biologist Laboratory de Physiologie, Val de Grace, Paris, 1952-1953, 56-61. Retired, 1961; visiting scientist International Cooperation Administration, fellow National Institutes of Health, 1954-1956. Director research department Rhone Poulenc Laboratories, Vitry-sur-Seine, 1961-1966.
Research group director Institute National de la Sante et de la Recherche Medicale, Le Vesinet, since 1967. Science council International Society Research and Civilization Diseases and Environmental, Luxembourg, since 1977.