Came to the United States, 1936, naturalized, 1941. Guest investigator University of Michigan, 1936-1937. Research associate department zoology University of California at Berkeley, 1937-1942, associate department veterinary science Experiment Station College Agriculture, 1942-1948.
Medical bacteriologist, chief variation branch Chemical Corps Biological laboratories, Camp Detrick, Frederick, Maryland., 1948-1955. Professor microbiology Institute Microbiology, Rutgers University, New Brunswick, New Jersey, 1955-1972. Consultant United States Army Chemical Corps, 1955-1968.
United States Public Health Service, 1956-1972, unites states department of Defense, 1960-1968, National Science Foundation, 1967-1970. Board science counselors National Institute Allergy and Infectious Diseases, National Institutes of Health, 1966-1969, Chairman of the Board science counselors, 1968-1969, visiting scientist, 1970-1972. I.M. Lewis lecturer University Texas, 1951.
Visiting professor U. P.R. Medical School, 1957, 59, 60, 61, 63, 65, 67, 68. National Institutes of Health research fellow, Israel, Paris, also visiting professor Hebrew U., Hadassah Medical School, Jerusalem, 1962-1963. Visiting scientist Weizmann Institute, Israel, Pasteur Institute, Paris, Karolinska Institute, Stockholm, 1969-1970.
O. Stark lecturer Miami University, Oxford, O., 1971.
Fellow American Academy of Microbiology (president 1968-1969), American Association for the Advancement of Science. Member Theobald Smith Society (president 1957-1958), Science Research Society American, American Society Microbiology, American Association Immunologists, Genetics Society American, Society Experimental Biology and Medicine, Society General Microbiology, Sigma Xi, Phi Sigma.