United States Public Health Service fellow biochemistry New York University School Medicine, 1949-1952. Assistant professor, then associate professor microbiology Yale School Medicine, 1942-1961. Professor biology U. Rochester, 1961-1973, department chairman, 1965-1969.
Associate director Space Science Center, 1966-1971. CONS. National Aeronautics and Space Administration, National Academy Sciences. Chairman life science working group member Executive Committee COSPAR.
Lecturer Union of the Soviet Socialist Republics Academy Sciences, 1969. American Chemical Society lecturer, 1969, 72. Microbiology Foundation lecturer, 1972.
Delaware Connecticut Democratic Convs., 1958.