Party affiliation: Democrat
Research associate Massachusetts Institute of Technology, Cambridge, 1964-1966. Assistant professor University of California at Los Angeles, 1966-1967. Assistant professor microbiology University Vermont, Burlington, 1967-1971, associate professor, 1971-1974, professor, since 1974, chairman department, 1979-1988, associate dean Graduate College, since 1994.
Secretary Genetic Resources Inc., Burlington, since 1982.