Bachelor of Science, Lafayette College, Easton, Pennsylvania, 1923, Honorary Doctor of Science, 1959. Doctor of Medicine Johns Hopkins, 1927, honorary Doctor of Laws, 1969. Doctor of Science (honorary), University Pennsylvania, 1971.
M.D. (honorary), University Freiburg i/B, 1971. Doctor of Science (honorary), Rockefeller U., 1976, U. Maryland., 1978, Syracuse University, 1979. Eldridge Johnson traveling research scholar, U. Leipzig and Munich, 1929-1931.
Research fellow medical science Johns Hopkins, 1927-1929. Fellow in medical physics Eldridge Johnson Research Foundation, University of Pennsylvania, 1931-1936, assistant professor biophysics, 1936-1940, 41-42, associate professor biophysics, 1943-1948, professor, 1948-1949. Associate professor physiology Cornell University Medical College, New York City, 1940-1941.
Professor biophysics, department chairman Johns Hopkins, Baltimore, 1949-1953. Professor Rockefeller U., New York City, 1953-1974, emeritus, 1974-1983.
Born December 22, 1903