Technician Rockefeller Institute Medical Reserach, 1937-1939. Assistant biochemistry Cornell, 1939-1943. Boichemist Office of Scientific Research and Development, New York University, 1944, instructor physiology college medicine, 1945-1948, assistant professor, 1948-1951, associate professor, 1951-1952.
Professor physiology, department chairman, school medicine University of Virginia since 1952.
Born September 20, 1912