- Instructor anatomy University of Michigan, 1942-1946. Assistant professor of University Washington, 1946-1948, associate professor, 1948-1957, administrative officer, 1955-1961, professor, 1957-1978, department chairman anatomy, professor, department chairman biological structure, 1961-1978.
Member anatomical science training grant committee National Institutes of Health, 1963-1964, 69-73, chairman, 1971-1973. Member anatomy test committee National Board Medical Examiners, 1967-1972. Member American Association Anatomists (president 1977-1978), American Society Immunology, American, International societies cell biology, International, American societies hematology, Royal Society Medicine (affiliate), Reticuloendothelial Society (councillor 1972-1976, president-elect 1978), Association Anatomy Chairman (1st president 1969-1970), Sigma Xi.