- Scientific assistant Max Planck Institute for Brain Research, Frankfurt, 1964-1968, 70-77. Professor ear, nose & throat clinic University Freiburg, head unit for morphological brain research. Visiting scientist Yerkes Regional Primate Research Center, Atlanta, 1969.
- Contributor articles to professional journals including Acta Anatomica, Behavioural Brain Research, Brain Research, Experimental Brain Research, Folia Primatologia, Hearing Research, Journal Comparative Neurology, Morphologisches Jahrbuch, Transactions of the Royal Society Edinburgh, Neurosci. Letters.