Instructor Duke University, 1957-1958, faculty fellow, Southern Fellowship Fund fellow, 1958-1959. Member faculty University Rhode Island, 1959-1970, professor biology, 1968-1970. Director Institute Environmental Biology, 1966—1970.
Professor, chairman department zoology University Oklahoma, Norman, 1970-1980, George Lynn Cross research professor zoology, 1979-2001, research professor emeritus, since 2001. Research professor Universidad de Los Andes, Bogotá, Colombia, 1965-1966. Principal investigator National Geography Society-U. Rhode Island herpetological expedition to Colombia, 1964-1965, National Geography Society-U. Oklahoma expeditions to Lake Titicaca, 1975, Cameroon, 1981.
With United States Army, 1952-1954, colonel medical service corporation United States Army Reserve.
- Editor Animal Natural History series, since 1991. Research and articles on heat tolerances of lower vertebrates, effects of day-length on metabolism and temperature tolerance of lower vertebrates, physiology of lower vertebrates, physiological ecology of amphibians and reptiles, respiration in amphibians, behavioral thermoregulation.
Fellow American Association for the Advancement of Science. Member American Institute Biological Science, American Society Ichthyologists and Herpetologists (president 1988), American Physiological Society, Ecological Society of America, Herpetologists League (executive committee 1968-1971), Society Study Amphibians and Reptiles (board governors 1986-1988, president 1998-1999), Explorers Club, Sigma Xi, Phi Sigma, Phi Kappa Phi, Oklahomans for Excellence in Science Education(founder, 2002).
1966 - 1970
1970 - 1980
1970 - 1980
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