- Instructor University Minnesota, Minneapolis, St. Paul, 1958-1959, assistant professor, 1959-1964, associate professor, 1964-1971, assistant dean College Biological Sciences, 1966-1969, associate dean College Biological Sciences, 1969-1979, professor, since 1971.
Member advisory committee University Research Consortium, Minneapolis, 1984-1991, Senate Consultant Committee, 1989-1992, vice chair, 1991-1992. Fellow American Association for the Advancement of Science. Member American Institute Biological Science, American Society Zoologists, Society for Development Biology, American Microscopical Society, Minnesota Academy of Sciences, American Association of University Professors (president University Minnesota chapter 1983-1984).
1958 - 1959
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