Bachelor of Arts, University Illinois, 1923. Master of Arts, Columbia, 1927, Doctor of Philosophy, 1929.
Assistant in psychology Barnard College, Columbia, 1927-1928. University fellow Columbia, 1928-1929. Research associate Yale, 1929-1933, associate professor psychobiology, 1933-1939, associate professor, 1939-1944, research associate, 1944-1956.
Assistant director Yerkes laboratories of Primate Biology, 1939-1952, associate director, 1952-1955, director, since 1955. Professor psycholobiology, Emory University, since 1956. Research associate Psychiatric Institute and Hospital, New York City, 1944, research consultant, 1946-1947.
Member American (division experimental psychological), Florida (president 1954-1955) psychological association, National Academy Sciences, Society Experimental Psyhcologists, American Association for the Advancement of Science, American Society Naturalists.
Married June 11, 1927. Children: Dora Jane, Joanna Marie.