Student Cornell Univercity, summer 1906. Bachelor of Arts, University of Michigan, 1908. Doctor of Philosophy, Columbia, 1911.
Entomological Michigan Biological Survey, summer, 1908. Instructor zoology, 1911, assistant professor, 1912, associate professor, 1914, professor since 1921, University of Michigan. Visiting professor University of California.
Fellow of the American Association for the Advancement of Science, Entomological Society American. Member American Society Zoölogists, American Society Naturalists (secretary 1920-1926, vice president 1929, president 1934), Genetics Society, Society for Study Evolution, American Genetic Association, Michigan Academy Science (president 1921-1922), Sigma Xi, Gamma Alpha. Henry Russel lecturer, 1951.
Married Margaret Jeffrey Buckley, December 23, 1911. Children: William Buckley (deceased), Catharine Ryman (deceased), Elizabeth Buckley (Mistress Elizabeth Russell), Franklin Buckley.