Came to the United States, 1898, derivative citizenship. Bachelor of Arts, Oberlin (O) College, 1904. Master of Arts, Harvard, 1905; student Strassburg (Alsace) University, 1906-1908.
Doctor of Philosophy summa cum laude, Wuerzburg (Germany) University, 1910.
Fellow in biochemistry, Toronto University, 1910-1911. Instructor physiology, University of Minnesota, 1911-1912, University of Missouri, 1912-1913, assistant professor, 1913-1918, associate professor, 1918-1921, professor physiological chemical University Missouri Medical School since 1921, chairman biochemistry department since 1931.
Fellow in biochemistry, Toronto University, 1910-1911. Member American Association for the Advancement of Science, Missouri Academy Science, American Chemical Society American Zoölogists, American Physiological Society, Society Study Internal Secretions, American Society Naturalists, American Institute Nutrition, Sigma Xi. Club: Appalachian Mountain.
Married Margaret Buckingham, March 29, 1916. Children: Mary Buckingham, Anne Darracott.