Doctor of Philosophy, University Nottingham, England, 1960. Doctor of Science, University Nottingham, England, 1973. Doctor of Laws (honorary), University Nottingham, England, 2005.
Research associate Indiana University, Bloomington, 1960-1962. Assistant professor chemistry Case Western Reserve University, Cleveland, 1962-1966, associate professor, 1966-1969, associate professor macromolecular science, 1969-1971, professor, 1971-1981, director laboratory for biological macromolecules, 1972-1981, distinguished adjunct university professor, since 2007. President, Chief Executive Officer University Genetics Company, 1981-1986, chairman, 1986-1987.
Senior partner Oxford Biosci Partners, Westport, Connecticut, since 1987. Chairman Oxford Bioscience Corporation, since 1992. President Committee Business Innovation, since 2007.
Visiting lecturer biological chemistry Harvard Medical School, 1971—1972. Member President Carter's Task Force on Science and Technology. United States project officer Rudjer Boskovic Institute, Zagreb, Yugoslavia, 1967—1975.
Board directors Acadia Pharma, Alexandria R.E.I.T. Emeritus director Research!America. Founder, past chairman Gene Logic, Inc., Exelixis, Inc.
Co-founder, past board member Human Genome Sciences, Inc. Co-founder The Institute for Genomic Research, 1994, National Committee Bioscis. Innovation, since 2008, president, since 2008.
Member and communications advisory President Obama's Policy Committee Science and Technology, 2008—2009.
Board directors Friends of Nottingham University. Member National Venture Capital Association, Sigma Xi (Research award 1973), Pi Kappa Alpha.
Married Jasmin Yvonne Christensen, September 1, 1958 (deceased November 1970). Children: Kimm A., Keir D.A. Married Elenor Jean McElliott, August 6, 1977.
Children: Kristin M., Sherri L.