Bachelor, California State University, 1971; Master of Arts, California State University, 1973; Doctor of Philosophy, Duke U., 1978.
Assistant professor botany, U. Maryland., College Park, 1979-1982;
associate professor botany, U. Maryland., College Park, 1982-1988;
professor botany, U. Maryland., College Park, 1988-1993;
dean College Natural Sciences, U. Hawaii, Honolulu, since 1994;
senior vice president for research, dean graduate divsin., U. Hawaii, Honolulu, since 1997. Guest professor Botanishes Institute, Karlsruhe, Germany, 1982-1983. Chairman sci.adv. board Center Global Change, College Park, 1989-1993.
Consultant USDA, Environmental Protection Agency, National Academy Science, Washington, since 1982.
Member American Society Plant Physiologists, Botanical Society American, Ecological Society American, Sigma Xi.
Married Karen Lee McKnight, September 10, 1974. 2 children.