Master of Arts, Aberdeen (Scotland) University, 1933. Bachelor of Science, Aberdeen (Scotland) University, 1936. Bachelor of Medicine ChB, Edinburgh University, 1940.
Doctor of Medicine, Edinburgh University, 1945. Doctor of Science (honorary), St. Andrews University, Scotland, 1972. Doctor of Laws (honorary), Strathclyde, Glasgow, 1985.
Carnegie scholar University Edinburgh, 1940-1941, lecturer in physiology, 1941-1948. Professor physiology University St. Andrews, 1948-1969. Secretary, treasurer Carnegie Trust for Universities of Scotland, 1969-1986.
Chairman Government and Educational Bodies, Scottish Hospital Endowment Trust, 1967-1969. Consultant in field.
Senior science advisor Civil Defense, Edinburgh, 1961-1980. Member British Library. Board, 1973-1980; trustee The National Library Scotland, 1975-1995.
Fellow Chartered Society Physiology, Royal College Physicians of Edinburgh (honorary), Royal Society Edinburgh (general secretary, vice president since 1953, Bicentenary medal 1983). Member British Medical Association.
Married Elizabeth Lambie Knox, June 28, 1941. Children: Innes, James, Margaret, Alison.