Bachelor of Science, Sussex University, Falmer, United Kingdom, 1980. Doctor of Philosophy, Oxford (United Kingdom) University, 1985.
Research associate department botany, Louisiana State University, Baton Rouge, 1983-1987; research fellow, Rutgers University, New Brunswick, New Jersey, 1987-1989; principal investigator, Dupont Experimental Station, Wilmington, Delaware, since 1989.
Member American Society Biochemistry and Molecular Biology, American Society Plant Physiologists, Sigma Xi.
Married Alison Jo Mack.