Grad, University Tübingen, 1967. Doctor of Philosophy, University Göttingen, Munich, 1974. Degree (honorary), University Alicante, Spain.
Degree (honorary), University Liverpool. Degree (honorary), University Bordeaux, France. Degree (honorary), University Munich, Germany.
Degree (honorary), University Colorado, Denver. Degree (honorary), University College London. Degree (honorary), Weizmann Institute, Rehovot, Israel.
Research assistant department of neurophysiology Max Planck Institute for Psychiatry, Munich, 1969—1970. Council fellow department of biophysics University College, London, 1971—1973. Research assistant department of neurobiology Max Planck Institute for Biophysical Chemical, Göttingen, Germany, 1974—1979, research associate membrane biology group Germany, 1979—1982, director membrane biology group Germany, 1983—1985, director department of cell physiology Germany, 1985—1989, Max Planck Institute for Medical Research, Heidelberg, Germany, since 1989.
Foreign member National Academy of Sciences, Royal Society, London.
Member of Orden Pour le Mérite, Heidelberger Academy of Sciences, Goettinger Academy of Sciences, Bavarian Academy of Sciences and Humanities, German Academy of Natural Scientists Leopoldina (Carus medal 1991).