Bachelor of Science, Reading University, England, 1952. Doctor of Philosophy, London University, 1955. Digital Array Gas-cell correlation radiometer and Forestry (honorary), Helsinki (Finland) University, 1990.
Research trainee Rothamsted Experimental Station, Harpenden, England, 1952-1954, senior science officer England, 1957-1966. Science officer Scottish Horticultural Research Institute, Dundee, Scotland, 1954-1957. Principal, senior principal then deputy chief science officer Scot.
Crop (formerly Horticultural) Research Institute, 1966-1991. Professor Dundee University, 1991-1996, professor emeritus, since 1997. Visiting scientist Waite Agricultural Research Institute, Adelaide, Australia, 1964.
Honorary professor University St. Andrews, Scotland, since 1986. Behrens lecturer University Leeds, 1990.
Fellow Royal Society London, Royal Society Edinburgh. Member Association Applied Biologists (honorary), Society General Microbiology (honorary), Phytopath. Society Japan (honorary).
Married Elizabeth Ann Latham-Warde, January 13, 1968. Children: Peter, Robert, Claire.