Student, U.'s Tübingen and Hamburg, Federal Republic of Germany. Doctor of Medicine, University Kiel, Federal Republic of Germany, 1965. Doctor of Philosophy in medicine and physiology, University Kiel, Federal Republic of Germany, 1968.
Habilitation, University Kiel, Federal Republic of Germany, 1969.
Professor physiology, director electrophysiology Department, University Bochum, Federal Republic of Germany, 1970-1973; professor, director neurophysiology Department, University Hamburg, since 1974; head Institute of Physiology, U. Hamburg, since 1977.
Member International Association Study of Pain (local organizer world congress Hamburg 1987), European Association Neurosci. (co-organizer congress in Hamburg 1983), German Society Physiology and Pharmacology, International Society Physiology and Pharmacology.
Married Thussi Klemm. Children: Karen, Boris.