Bachelor of Science, Iowa State University, 1963. Master of Science, Illinois Institute of Technology, 1968. Doctor of Philosophy, Indiana University, 1971.
Research biochemist Illinois Institute of Technology Research Institute, Chicago, 1963-1968. Graduate fellow Indiana University, Bloomington, 1968-1971. Postdoctoral fellow Cornell University, Ithaca, New York, 1971-1973.
Assistant professor University Michigan, Ann Arbor, 1973-1977, associate professor, 1978-1982, professor biological science and chemistry, since 1983. Alfred S. Sussman distinguished university professor, since 2004, chairman department biology, 1985-1991. Visiting professor Michigan State University, East Lansing, 1980-1981.
Consultant National Science Foundation, Washington, since 1982.
Fellow American Association for the Advancement of Science. Member American Chemical Society, American Society Plant Physiologists, American Society Biological Chemists, American Society Photobiology.
Married Patricia Joan Bury, January 1, 1981. 1 son, Erik Charles.