Bachelor, Cambridge University, 1944. Doctor of Philosophy, Aberdeen University, 1948. Doctor of Science, Cambridge University, 1977.
Research assistant Imperial College, London, 1944-1945. From assistant lecturer botany to professor botany University Aberdeen, United Kingdom, 1945-1988. Head department botany University Aberdeen, 1981-1988.
Scottish advisory committee Nature Conservancy Council, 1966-1980. Member Countryside Commission for Scotland, 1980-1991.
Active Northeast Regional Board, Scottish Natural Heritage, since 1992.
Married Elizabeth Caroline Baird, August 21, 1948. Children: Alison Caroline, Anne Fiona, Clare Elizabeth.