Bachelor of Science, University of California, Davis, 1957; Master of Science, University of California, Davis, 1959; Doctor of Philosophy, University of California, Davis, 1964.
Assistant professor plant pathology University Florida, Gainesville, 1964-1969, associate professor, 1969-1975, professor, 1975-1999, professor emeritus, since 2000. Retired, 2000
Plant virus subcommittee International Committee for Taxonomy of Viruses, 1973-1975, member potyvirus study group, 1987-1993. Member plant virology advisory committee American Type Culture Collection, 1993-1999.
Member International Legume Virus Working Group, 1999. With United States Army, 1957.
Member Morningside Nature Center Commission, City of Gainesville, 1978-1981, treasurer, 1981. Fellow American Association for the Advancement of Science, American Phytopathol Society (Lee Hutchins award 1981, Ruth Allen award 1992). Member Florida State Horticultural Society, New York Academy of Sciences, American Society Virology, Phytopathol.
Society Japan, Australasian Plant Pathology Society, Brazilian Phytopath. Society, United States Golf Association, National Wildlife Federation (associate), Nature Conservancy, National Geographic Society, Smithsonian Institution (associate), Sigma Xi, Gamma Sigma Delta.
Married Marcia Ann Weatherby, September 7, 1966. Children: Scott, Douglas.