Bachelor, New York University, 1970. Doctor of Philosophy, State University of New York Health Science Center, Brooklyn, 1978.
Assistant research scientist department cell biology, New York University School Medicine, New York City, 1978-1983; assistant professor department cell biology, New York University School Medicine, New York City, 1983-1987; associate professor departments anatomy, cell biology, pathology, College Physicians and Surgeons Columbia University, New York City, since 1987.
Member Harvey Society, New York Academy of Sciences, American Society Cell Biology, Society Neurosci.
Married Elizabeth Ann Wikstrom.