Doctor of Veterinary Medicine, National Veterinary School of Nantes, France, 1988. Doctor of Science, Paul Sabatier University, Toulouse, France, 1990.
Assistant National Veterinary School of Nantes, 1986-1987. Research assistant National Institute for Agronomic Research, Toulouse, 1988-1990. Research associate Lund (Sweden) University, 1990-1993.
Established investigator National Center for Science Research (National Center for Scientific Research), Toulouse, since 1993.