Bachelor of Science cum laude, Syracuse University, 1959. Master of Science, Syracuse University, 1962. Doctor of Philosophy, Syracuse University, 1968.
Research assistant zoology Syracuse University, 1957-1962. Teacher biology and general science Baker High School, Baldwinsville, New York, 1960-1962. Teaching assistant zoology Syracuse University, 1962.
Research associate microbiology State University of New York, 1963-1964. United States Public Health Service trainee in comparative physiology, zoology Syracuse University, 1963-1967. United States Public Health Service postdoctoral fellow in cell biology University Chicago, 1968-1970.
Postdoctoral fellow in biochemistry, biology Bryn Mawr (Pennsylvania) College, 1970-1972. Assistant professor to associate professor department biological science University Cincinnati, 1972-1982, professor department biological science, since 1982. Senior research microbiologist Laboratory Parasitic Diseases National Institute Allergy and Infectious Diseases, National Institutes of Health, Bethesda, Maryland, 1980-1981.
Member cell biology advisory panel National Science Foundation, 1983-1986, biochemistry, biophysics and molecular biology panel, since 1987, chair, since 1989. Member ad hoc study sections National Institutes of Health, 1974, 82, 89.
Member American Association for the Advancement of Science, American Association of University Professors, Society Protozoologists (nominating committee 1982-1983, consultant 1983, chair program committee 1984-1988, workshop organizer, 1988, conference organizer 1989, ), American Society Cell Biology (chair local arrangements committee 1980, program committee 1980, membership committee 1981-1982, legislation alert committee since 1981), American Women in Science, University Cincinnati Association for Women Faculty (executive board 1974-1975), New York Academy of Sciences, American Society Tropical Medicine and Hygiene, Marine Biological Laboratory (member corporation since 1973), Sigma Xi (chapter secretary 1987-1989, vice president 1989-1991).