Bachelor of Science, Cornell Univercity, 1954; Master of Science, North Carolina State College, 1956; Doctor of Philosophy, Columbia University, 1964.
Instructor department mathematics and astronomy, Lehigh University, Bethlehem, Pennsylvania, 1963-1964; assistant professor department statistics, Iowa State University, Ames, 1964-1967; associate professor, Iowa State University, Ames, 1967-1972; professor statistics, Iowa State University, Ames, since 1972.
With United States Army, 1956-1958. Fellow American Statistical Association. Member American Society Naturalists, Genetics Society American, Biometric Society.