Bachelor of Arts in Biology, Stanford University, 1963. Doctor of Philosophy in Bio-chemical, University Washington, 1968.
Professor biochemistry University California, Berkeley, 1971—1981. Co-founder, Chief Executive Officer Chiron Corporation, 1981—1998. Dean School Public Health University California, Berkeley, 1998—2002, dean emeritus, since 2002.
Senior director, Science & Higher Education Gordon and Betty Moore Foundation, 2002—2004, president, 2004—2008, board trustees. Board directors, senior advisory to Chief Executive Officer Chiron Corporation.
Member of American Society of Biological Chemists, National Academy of Science, Institute Medicine.