Bachelor of Science, University of Minnesota, 1959, Master of Science, 1966. Doctor of Philosophy, University of Wisconsin, 1970. Certified wildlife biologist.
Assistant professor, zoology and veterinary science North Dakota State University, Fargo, 1968—1971. Assistant leader, Oregon cooperative wildlife research unit United States Fish and Wildlife Service, Oregon State University, Corvallis, 1971—1976, leader, since 1976. Lieutenant (junior grade) United States Navy, 1959-1962.
Served to Lieutenant (junior grade) United States Navy, 1959-1962. Member Wildlife Society (section representative 1977-1980, vice president 81-83, president 1984-1986), Ecological Society American, American Society Mammalogists, American Ornithologists Union, International Association.Bear Research and Management.
Married Elaine Frances Stone, December 19, 1959. Children: Karen, Kirstin, Eric.