Master of Science in Microbiology, Tel Aviv University, 1964. Doctor of Philosophy in Genetics, Weizmann Institute of Science, Rehovot, Israel, 1969. Doctor honoris causa, Russian Academy of Sciences, Moscow, 1997.
Visiting scientist National Cancer Institute, 1968, 71. Senior scientist, associate professor Weizmann Institute of Science, Rehovot, Israel, 1971, 77. Group leader, senior scientist Oak Ridge (Tennessee) National Laboratory, 1976-1981.
Professor University Chicago, 1982-1997. Group leader, senior biologist Argonne (Illinois) National Laboratory, since 1981. Visiting professor Kobe University, 2000.
Member American Association Cancer Research, Radiation Research Society, New York Academy of Sciences.
Married Lily Ginzburg, May 11, 1967. Children: Ilan, Ron.