Bachelor of Science, University North Dakota, 1968. Master of Science, Southern Illinois University, 1972. Doctor of Philosophy, University Minnesota, 1976.
Research assistant, Delta Waterfowl and Wetlands Research Station, Portage la Prairie, Manitoba, Canada, 1968-1971; research assistant, Northern Prairie Wildlife Research Center, Jamestown, North Dakota, 1972-1977; teaching assistant, University of Minnesota, St. Paul, 1973-1976; instructor, McGill University, Montreal, Quebec, Canada, 1976-1977; from. assistant professor to professor School Natural Resource, U. Missouri, Columbia, since 1978; program leader, U. Missouri, Columbia, since 1989. Coordinator Missouri U. Wetland Center, Columbia, since 1991. Director Ashland Wildlife Research Center, Columbia, 1980-1988.
Member Wildlife Society (council representative 1986-1989, vice president since 1992, president North Center section 1991-1992, president Missouri chapter, president-elect 1993-1994), American Society Mammalogists, Society Conservation Biologists.
Married Ell-Piret Multer, July 15, 1973. Children: Leif, Helen.