Bachelor of Science, Liverpool (England) University, 1964. Bachelor of Science with honors, Liverpool (England) University, 1965. Doctor of Philosophy, Liverpool (England) University, 1968.
Postdoctoral research fellow Birmingham University, 1968-1970. Lecturer virology Queen's University, Belfast, Northern Ireland, 1970-1979. Senior virology lecturer London University, 1979-1988.
Assistant director Institute Virology, Oxford, England, since 1988. Visiting professor University Tripoli, 1980-1990. Member Health and Safety Executive advisory committee Dangerous Pathogens, since 1992.
Member American Society Virology, Society General Microbiology.
Married Suzanne Mary Holliday, August 20, 1966. Children: Kathryn Louise, Richard Andrew, Jonathan Edward.