Master of Public Health in Plant Physiology, University Turku, Finland, 1987. Doctor Plant Physiology, University Turku, Finland, 1993.
Researcher photosynthesis, U. Turku, 1987-1994, 96-; docent in plant physiology and biophysics, U. Turku, since 1995; managing director, Quaid-i-Azam-Data Oy, since 1995.
Member Scandinavian Society Plant Physiology (treasurer since 1997).
Married Eija Koivukoski, 1981 (divorced 1992). Children: Joonas, Maria, Venla. Married Taina Kallio, 1993.
Children: Sofia, Topias.