Education
Complutense University of Madrid.
Complutense University of Madrid.
He was also keenly interested in the historical biogeography of these groups. He conducted research in community ecology with the idea of trying to ascertain the underlying causes that determined the diversity of dung beetle communities. He was the coordinator for the Insecta portion of the Fauna Iberica project and is the author of the first volume on the Scarabaeoidea of the Iberian Peninsula.