Student, University Tübingen, Federal Republic Germany, 1954. Doctors degree, University Göttingen, Federal Republic Germany, 1962.
Researcher Max Planck Institute for Cell Biology, Tübingen, 1962-1966. Researcher, lecturer Zoological Institute of the Un., Würzburg, Federal Republic Germany, 1966-1970, assistant professor, 1971-1972, associate professor, 1972-1977. Visiting researcher Institute Genetics University Lund, Sweden, 1963, Department Zoological University St. Andrews, Scotland, 1967.
Full professor, director Zoological Institute of the University, Kiel, Germany, 1977—1998. Retired, 1998
Member advisory board Scientific Journal Entomologia Generalis.
Member German Zoological Society, Society for Genetics, Society for Developmental Biology.
Married Rosemarie Borck, September 21, 1962. Children: Hansjörg, Stefan.