Bachelor of Science in Zoology, Oregon State University, 1973. Doctor of Philosophy in Genetics, University Washington, 1977.
Postdoctoral researcher, University of California, Davis, 1978-1979; assistant professor, Washington State University, Pullman, 1979-1986; associate professor, Washington State University, 1986-1990; professor, Washington State University, since 1990. Chair department zoology Washington State University, since 1991. Co-organizer North Atlantic Treaty Organization Conference onGenetic Conservation of Salmonid Fishes, 1991.
Fellow American Association for the Advancement of Science. Member American Fisheries Society, Genetics Society of America, International Association Genetics in Aquaculture.
Married Beverly J. Hunter, August 20, 1983 (divorced January 1995).