Bachelor of Science, Iowa State University, 1969; Doctor of Philosophy, Stanford University, 1974.
Postdoctoral, U. Basel, Switzerland, 1974-1977; postdoctoral, University of California San Diego, La Jolla, 1977-1978; assistant professor department biology, Boston University, 1979-1985; associate professor, Boston University, 1985-1991; professor, Boston University, since 1991.
Fellow American Association for the Advancement of Science. Member American Society Microbiology, American Society Biochemistry and Molecular Biology.
Married Paula Lillian Grisafi, September 29, 1984.