Bachelor magna cum laude, University of Pennsylvania, 1983; Master of Science, Yale University, 1984; Doctor of Philosophy, Princeton University, 1989.
Postdoctoral fellow New York University Medical Center American Cancer Society, 1989-1991. Senior research biochemist American Cancer Society Merck and Company, Inc., West Point, Pennsylvania, 1991-1993. Assistant professor Wistar Institute, University Pennsylvania, Philadelphia, 1993-1997, associate professor, 1997—2001.
Senior director cancer research DuPont Pharmaceuticals Company, Glenolden, 1999—2001. Professor Lankenau Institute Medical Research, since 2002, president, Chief Executive Officer, since 2004. Professor Thomas Jefferson University, Philadelphia, since 2002.
Member American Association Cancer Research, American Cancer Society.
Married Kristine Kushmeider, October 25, 1986.