Bachelor of Science, Novosibirsk State University, Russia, 1974. Doctor of Philosophy, Institute Cytology, Novosibirsk, 1980. Doctor of Science, Institute Cytology & Genetics, 1987.
Head of laboratory Institute Cytology & Genetics, Novosibirsk, 1986—1990. Senior research fellow Imperial Cancer Research Fund, Clare Hall Laboratories, South Mimms, 1990—1993. Senior staff fellow National Institute on Aging, National Institutes of Health, Baltimore, 1995—2000.
Senior scientist Medical Research Council Radiation & Genome Stability Unit, Harwell, England, since 2000.
Member of American Society Biochemistry and Molecular Biology, American Radiation Research Society.
Son of Herman Davidovich Kofler (Stepfather) and Eugenia Petrovna Dianova. Married Irina Iosifovna Kolker, November 30, 1972. Children: Anton Grigorievich, Vera Grigorievna.