Staatsexamen, University Mainz, Germany, 1957. Doctor Natural Sciences, University Mainz, 1958.
Scholar, German Research Association, Braunschweig, Germany and Naples, Italy scholar, German Research Association, Banyuls, France scholar, German Research Association, Kuala Lumpur, Malaysia scholar, German Research Association, Bogor, Indonesia, 1958-1962; research officer, Food and Agriculture Organization (of the United Nations), Banagi-Serengeti, Tanzania, 1962-1965; research assistant, U. Brunswick, 1965-1972; professor zoology, U. Brunswick, since 1972. Chairman Equidae group International Union for Conservation of Nature and Natural Resources-Species Survival Commission, 1977-1987, member, since 1987, member Hippopotamidae group, since 1981.
Member American Association Mammalogists, Deutsche Zoologische Gesellschaft, Deutsche Gesellschaft für Säugetierkunde, East African Wildlife Society, Fauna and Flora Preservation Society, Frankfurt Zoological Society, also others.
Married Ute Sievers, September 17, 1963.