AB, Kenyon College, 1962. Doctor of Science (honorary), Kenyon College, 1982. Doctor of Philosophy, Rockefeller University, 1966.
Postdoctoral fellow Medical Research Council Laboratory of Molecular Biology, Cambridge, England, 1966-1968. Assistant professor biology Massachusetts Institute of Technology, Cambridge, Massachusetts, 1968-1971, associate professor, 1971-1976, professor, 1976. Vis.scientist Imperial Cancer Research Fund, Lincoln"s Inn Fields, London, 1977-1978.
Visiting professor University of California, San Francisco Medical Center, 1977, Pennsylvania State University Medical Center, 1977.
Staples visiting professor U. Maine, 1989. Wellcome visiting professor U. Oregon, 1990.
Roering visiting lecturer U. Washington Medical School, 1991. Sackler visiting scholar Tel Aviv University, 1994.
Berson Memorial lecturer International Endocrinology Congress, 1992, Presidential lecturer International Hematology Congress, 1992, McGinnis Memorial lecturer, 1994, Sigma Xi lecturer, 1994, Harry Eagle Memorial lecturer, 1994.
Consultant scientist medicine Children"s Hospital Medical Center, Boston, 1977. Consultant in pediatric oncology Sidney Farber Cancer Institute, 1977. Consultant Ministry Edition and Research, Denmark, 1991-1992.
Member Whitehead Institute biomedical Research.
Science advisory board Damon Biotechnology, Incorporated., Fisher Science Group, Genzyme, Incorporated., Arris Pharmaceutical Company, Millennium Corporation, Astra Bachelor of Arts, Sweden, Cogito, Incorporated. President Bioinfo. Associations, Incorporated.
Consultant biotechnology. Member National Science Foundation advisory panel in development and cellular biology;member National Institutes of Health study section of molecular biology and cell biology.
Advisory bd.Biozentrum, U. Basel, Switzerland, Fred Hutchinson Cancer Center, chair, Seattle, Cleveland Clinical Research Foundation, chair, European Molecular Biology Laboratory, Heidelberg, Germany, Center for Molecular Biology, Heidelberg.
Consultant National Institutes of Health, American Cancer Society Vice chairperson Gordon Research Conference, 1975, chairperson, 1976, 85, 89. Trustee Kenyon College, 1989.
Member Whitehead Institute biomedical Research. Member National Science Foundation advisory panel in development and cellular biology. Fellow American Association for the Advancement of Science, American Academy Microbiology.
Member National Academy of Sciences (member editorial board proceedings 1996), American Society for Microbiology, American Society of Biological Chemists (nominating committee 1979, 86), American Society for Cell Biology (county 1983-1985), American Chemical Society, European Molecular Biological Organisation (associate foreign), Phi Beta Kappa, Sigma Xi.
Married Pamela Chentow, December 29, 1963. Children: Heidi, Martin, Stephanie.