Doctor, University Ljubljana, Slovenia, 1963. Doctor, Goteborg University, Sweden, 1963.
Assistant Biological Institute, Medical Faculty U., Ljubljana, Slovenia, 1955-1960; research fellow, Norwegian Institute Algological Research, Trondheim, 1960-1963; scientific official, Hydrobiologish Institute, Yerseke, Holland, 1963-1965; research assistant Institute Food Technology, Ljubljana (Slovenia) U., 1965-1966; associate researcher Biological Institute, Slovene Academy Science and Arts, Ljubljana, 1966-1983; scienific official Center Scientific Research, Slovene Academy Science and Arts, Ljubljana, since 1983.
Member British Phycol. Society, International Phycol. Society, Phycol. Society of America, New York Academy of Sciences, International Commission Scientific Exploration of the Mediterranean Sea (CIESM) (Monaco), European Cooperation in Field of Scientific and Technology Research (Cooperation in Science and Technology), German Botanical Society, The Planetary Society, International Governors Club.