Bachelor of Science in Chemistry, Montana State College, 1958. Master of Science in Biochemistry, University North Dakota, 1960. Doctor of Philosophy in Biochemistry, University North Dakota, 1970.
Chemist, National Institutes of Health, Bethesda, Maryland., 1961-1969; research chemist, National Institutes of Health, Bethesda, since 1970.
Board of directors, secretary The Ivymount School, Rockville, Maryland., since 1985. Member American Chemical Society, American Society for Microbiology, American Society for Biochemistry and Molecular Biology, Sigma Xi.
Married Annette Marlane Dapp. Children: Janice, Susanne, Todd.