Education
He was educated at Winchester College.
He was educated at Winchester College.
He became professor of biological diversity at the Department of Biology, University College London. He was co-director of the Centre for Ecology and Evolution, a centre of excellence in research and teaching formed by University College London, the Institute of Zoology (Zoological Society of London), Natural History Museum, Imperial College, Queen Mary, Royal Holloway and Kew Gardens. In 2013 he was distinguished lecturer on Organismic and Evolutionary Biology at Harvard University.
He was awarded the Darwinby the Linnean Society of London in 2008.