Bachelor of Science, Yale University, 1963. Doctor of Philosophy, California Institute of Technology, 1967.
Chief biochemist, United States Army Aeromed. Research Lab, Fort Rucker, Alabama, 1967-1969; assistant professor of chemistry and biochemistry, University of Illinois, Urbana, 1969-1974; associate professor microbiology, Mount Sinai School Medicine, New York City, 1974-1982; professor, Mount Sinai School Medicine, New York City, since 1982. Consultant National Institutes of Health study section, Bethesda, Maryland., since 1987, Enzo Biochem, New York City, 1983-1992, Roche Molecular Sys., since 1992.
Active Town Club, Scarsdale, New York, since 1975. Captain United States Army, 1967-1969. Fellow New York Academy of Scientists (vice president 1986-1988).
Member American Society for Biochemistry and Molecular Biology, New York Science Policy Association.
Married Brigid Mary Long, September 4, 1965. Children: Katherine, John, Tara.