Bachelor, University of Virginia, 1973; Doctor of Philosophy, Yale University, 1979.
Postdoctoral fellow Ecosystems Center, Marine Biological Laboratory, Woods Hole, Massachusetts, 1979-1981. From assistant professor to associate professor department biology University South Carolina, Columbia, 1981-1993, professor department biology, since 1993. Faculty Water Resources division United States Geological Survey, 1988-1992.
Visiting professor department plant ecology Aarhus (Denmark) University, 1993. Consultant Environmental Protection Agency, since 1981. Member grant review panel United States Geological Survey, 1991, National Science Foundation, 1993-1998.
Reviewer proposals National Science Foundation, Environmental Protection Agency, United States Department of Agriculture, since 1981.
Member American Society Limnology & Oceanography, American Association for the Advancement of Science, Estuarine Research Federation, Sigma Xi. M C.
Married Maureen Mahoney. Children: Erin Maedora, Lily Elizabeth.