Bachelor of Science cum laude, Ohio State University, 1959; Master of Science, University of Michigan, 1963; Doctor of Philosophy, University of Michigan, 1968.
Assistant professor department biology, Pennsylvania State University, University Park, 1972-1974; associate professor, Pennsylvania State University, 1974-1981; associate scientist department genetics, S.W. Foundation for biomedical Research, San Antonio, 1981-1985; acting chairman, S.W. Foundation for biomedical Research, San Antonio, 1993-1994; scientist, S.W. Foundation for biomedical Research, San Antonio, since 1985; associate professor department cellular and structural biology, University Texas Health Science Center, San Antonio, 1982-1986; professor, University Texas Health Science Center, since 1986. Member various federal advisory committees and editorial boards. Executive positions in numerous organizations.
Member medical advisory committee South Texas Regional Blood Bank, San Antonio, 1988-1992. Member American Heart Association (fellow council arteriosclerosis), American Society Human Genetics (life), International Genetic Epidemiology Society, Phi Beta Kappa, Sigma Xi.