Education
He was educated at Winchester School, and Trinity College, Cambridge. He graduated from University College Hospital medical school in 1940.
He was educated at Winchester School, and Trinity College, Cambridge. He graduated from University College Hospital medical school in 1940.
He was a houseman at the Hammersmith Hospital, and was the Jenner research student at the Lister Institute, 1941-1942. He was assistant pathologist at the Central Middlesex Hospital from 1943-1946, then joined the external staff of the Medical Research Council as a bacteriologist at University College Hospital in 1946. joined the staff of the National Institute for Medical Research in 1949, working in the Division of Biological Standards. With James Lightbown he established international standards for antibiotics and enzymes, and later developed a long-standing association with the World Health Organization Expert Committee on Biological Standards.
In 1957 he became head of the Institute"s new Division of Immunology.
From 1961-1976 was Deputy Director of NIMR, and became acting director in 1969. In 1975, left NIMR to be Professor of Immunology at the Royal Postgraduate Medical School, Hammersmith.
He retired in 1981. In November 1956 founded the British Society for Immunology alongside Robin Coombs, Bob White, and Avrion Mitchison.
He was president of the International Union of Immunological Societies.
Royal Society.